Otolith morphology: Sagitta otolith fusiform, large in size, 14.9 and 15.4 mm in width respectively (Fig. 8A–
B). Distal surface concave, with three ridges on dorsal portion and one ridge on rostrum. Proximal surface slightly
convex; dorsal area of narrow, about same width with ventral area; dorsal depression and ventral depression absent;
crista superior not well-developed; dorsal margin lobed, with three lobes, the posterior lobe a short protraction;
ventral rounded in outline, with a crenate margin; anterior and posterior surfaces irregular. Excrisura a nearly right
angle gape; antirostrum less developed, with a flat or slightly convex margin; rostrum a large, pointed protraction
extending well beyond the antirostrum. Pseudo-excrisura a deep concave; pseudo-rostrum a large triangular
protraction. Sulcus groove very deep and wide, heteromorph; collum absent; a narrow and elongated ostium
(anterior sulcus) connected to the very wide and oval cauda (posterior sulcus) directly. Colliculum heteromorphy,
well-developed in cauda and less developed in ostium.