Nevertheless, at the breed level we observed substantial genetic differentiation, suggesting that in many cases, hybridization among breeds has been limited (mean pairwise (FST = 0.204) for all breeds, maximum (FST = 0.446); potentially more reliable differentiation estimates considering the modest sample sizes for some breeds [10]: mean( Dest = 0.15)6, maximum (Dest = 0.421); Tables S4 and S5).