Fruit coloration was measured by percent of fruit surface with red color and by
hue angle using a Minolta Chromameter (CR 200, Tokyo). Color values were
measured at both the reddest (exposed side) and greenest (shaded side) point of
the fruit's equator. Hue angle was calculated as described by McGuire (1992)
with 0 . red-purple, 90 . yellow, 180 . bluish-green, and 270 . blue.