In addition, tiger occupancy
was 12–30% greater than sites in Indonesia and India
(29, 30). Human foot traffic across the study site was also orders
of magnitude greater than traffic reported for other areas of the
tigers’ range (using similar methodology) (25, 26). Over the last
decade, tigers have maintained high densities in Chitwan (15,
31), although human density in settled areas surrounding the park
has increased 20% (212–255 people/km2) (32), approximately
two times the average human density (127 people/km2) among
12 of 13 tiger range countries (except Bangladesh) in 2010 (14).