Seasonal changes in day length (photoperiod) are consistent from year to year. Therefore, many plants use photoperiod information to predict upcoming environmental changes and precisely align the timing of flowering with favorable conditions [1]. Another important environmental factor that influences flowering is surrounding temperature. Ambient temperature changes arising from global climate change have already altered the phenology of plants, including the timing of flowering [2]. Therefore, understanding the mechanisms by which plants integrate both photoperiod and temperature cues to control seasonal flowering is necessary.