Leaves lose their lustre, green colour and normal curvature. Leaves appear leathery and curve downwards instead of the normal boat-shape; they discolour from deep-green to increasingly yellowish-brown. Initially, the discoloration of the foliage may affect only one branch, but it later spreads to the whole canopy. Infected trees may flower and fruit off-season. Eventually, leaves drop, branches die back and infected trees die, leaving large vacant spaces in severely attacked fields. On roots, networks of rhizomorphs are firmly attached. The growing ends of rhizomorphs form whitish fans. Mature rhizomorphs are brownish or may assume the colour of the surrounding soil. Severely infected roots are soft and watery with a creamy colour. Characteristic fruiting bodies form on tree collars.