In addition, we applied a Monte Carlo test of significance of observed
maximum indicator value, based on 1000 randomizations
(McCune and Grace, 2002). Only species with alpha levels less than
0.05 and with an indicator value greater than 25 were considered
indicator species for a treatment (Dufrêne and Legendre, 1997).
We applied a Bonferroni correction to account for multiple comparisons
in pairwise comparisons among treatments. All multivariate
tests were carried out using PC-ORD ver. 5 (McCune and
Mefford, 2006).