Wefound that a close coupling of the kinematics of the two oscillatingsystems appears to be precluded by their differential dependenceon the animal’s speed. In the case of the forelimbs, increases ordecreases in speed are mainly effected by changes in temporal gaitparameters such as stance phase duration and stride frequency.The frequency of whisk cycles, in contrast, varies independently ofspeed. On the other hand, forelimb position at touchdown is fairlyconstant relative to the head across the speed range, whereas thevibrissae change their protraction and retraction angles in accor-dance with speed.