ROS, especially H2O2, can diffuse across membranes and serve as regulatory molecules in signal transduction (Mubarakshina et al., 2010; Yamasaki et al., 1997). Under conditions of oxidative stress, the concentrations of some phenolic compounds, including nthocyanins, in plant tissue were shown to be greatly enhanced to protect the plants from oxidative damage (Zhang et al., 2010). This suggests that, besides anthocyanins, other phenolic compounds may also participate in ROS scavenging in vivo.