The ANOVA model included four main effects (land-use, soil depth, site, village nested in site) treated as fixed factors, and five two-way interactions (land-use by site, land-use by village nested in site, soil depth by site, soil depth by village nested in site, land-use by soil depth). The Tukey HSD (honestly significant difference) test was used to compare least-squares means for land-use systems, soil depths and sites.