They attach to solid surfaces in order to have access to nutrients in
the overlying liquid or from the solid matrix. Adhesion of bacteria
to surface is facilitated by exopolymeric substances, but hydrodynamic
conditions and the availability of substrate and nutrients
can shape biofilm architecture (Battin et al., 2007; Perdrial et al.,
2009). Physical and chemical interactions at bacteria-mineral
interfaces often determine the force of bacterial adhesion, and
are due to several factors