All these refugium-use strategies may be used by
animals during drought. The studies of Boulton
(1989) and Boulton et al. (1992c) of over-summering
strategies of macroinvertebrates undergoing seasonal
drought suggest that the most common strategies are
refugium use between generations through complex
life cycles and refugium use within generations and
within habitat. However, no comparative study has
been made of refugium-use strategies in streams
exposed to supra-seasonal drought, or has any study
examined the relative survival of fauna using different
types of refugia during a lengthy drought.