type. The pattern of change in response was
not symmetrical around 50:50 indicating that there is a complex
interaction between the farm type and its location that determineswhether the positive or negative effect is enhanced. The skylarkshowed a different pattern, whereby increasing heterogeneity bycombining farm types in the same landscape had a positive effectcompared to the arithmetic expectation, i.e. in this case hetero-geneity was always beneficial