Fiddler crabs (Uca spp.) occupy a wide range of habitats from salt marshes and
mangroves to sandy beaches (Crane, 1975). On beaches, they frequently live in the
intertidal zone where the sediment is subjected to cyclical exposure to air and submersion
under seawater. After the seawater recedes during low tide, fiddler crabs emerge
from their burrows to feed, attract mates, and defend their territory. One of the major
problems they face during emergence from their burrows is water loss through transpiration
via the carapace and gill surface, especially during feeding and excretion