The organism was observed in histological preparations to which nested polymerase chain reaction was used, with confirmation by sequencing . “Candidatus piscichlamydia salmonis” was detected by RT-DGGE in intracellular inclusions, i.e. epitheliocysts, in Atlantic salmon with proliferative gill inflammation . “Candidatus clavochlamydia salmonicola” is an intracellular organism, causing epitheliocystis in Atlantic salmon, which was recognized as novel as a result of 16S rRNA sequencing .