Thirty grams of fruit flesh tissue (derived from 5 fruit, with 3 replicates for each treatment) were homogenized using a FSH- 2A homogenizer. The homogenate was centrifuged at 8000 × g for 20 min at 4 ◦C and the supernatant was filtered through Miracloth. Two drops of the supernatant were dispensed onto a hand-held refractometer (Master-M, ATAGO Co. Ltd., Tokyo, Japan) to determine SSC. SSC was expressed as percentage. For TA measurement, 6 mL of the supernatant used for SSC were diluted with 50 mL of distilled water and the samples were titrated to pH 8.1 with 0.1 M NaOH. TA was expressed on a fresh basis as the percentage of citric acid equivalents per kilogram.