For the phylogenetic analysis, the 16S rRNA gene was amplified by PCR using the method described in Skillman et al. (2004) and the primers 21f (59-TTCCGGTTGATCCYGCCGGA-39) (DeLong, 1992) and 1386r (59-GCGGTGTGTGCAAGGAGC-39) (Skillman et al., 2004). The PCR products were cloned and then sent to MWG Biotech (Germany) for sequencing