Identification of agronomic important QTLs and pyramiding of such QTLs were presented
a useful strategy for efficient crop breeding. However, crop breeding is a complicated process.
In practice, many elite lines breeding experience revealed that breeding is a cumulative process
that includes many steps. In fact, other factors, such as grain quality, resistance to pests, and
other adversities, must also be taken into account. Besides growing duration and seed-setting
ability, numerous other crucial traits directly or indirectly relate to yield and quality. Many traits are