Tuberculation. In presumed fully developed males, numerous small tubercles cover the whole body, head and
parts of fins (Figs. 1B and E). Dentary tubercles on highly tuberculated males generally up to 5 incomplete rows in
large specimens, anterior ones larger than posterior ones, more distinct on outer and inner edges of dentary. Small
tubercles on side of snout, end of maxilla, front of eye, top of eye, all areas below eye, opercle and branchiostegal
region. Larger tubercles on entire side of body, generally up to 4 tubercles per scale in large specimens. Small and
large tubercles on snout, top of head, and back. Tubercles on presumed fully developed males also cover outer
surface of pectoral and pelvic fins, part of dorsal and anal fins, and the upper lobe of caudal fin. In normal
specimens tubercles restricted mainly to snout and dentary, and in these specimens they are not as numerous nor as
large as tubercles on presumed fully developed male.