-Conceptual diagram of an Integrated Multi-Trophic Aquaculture (IMTA) operation including the combination of fed aquaculture (e.g., finfish) with suspension organic extractive aquaculture (e.g., shellfish), taking advantage of the enrichment in small particulate organic matter (POM); inorganic extractive aquaculture (e.g., seaweeds), taking advantage of the enrichment in dissolved inorganic nutrients (DIN); and deposit organic extractive aquaculture (e.g., echinoids, holothuroids, and polychaetes), taking advantage of the enrichment in large particulate organic matter (POM) and feces and pseudo-feces (F&PF) from suspension-feeding organisms. The bioturbation on the bottom also regenerates some DIN, which becomes available to the seaweeds