Indeed, Atkrp125c mutants (also referred as radially swollen 7, rsw7)
exhibit massive spindle deformities and show spindle collapse (Wiedemeier et al., 2002; Bannigan et al., 2007), suggest- ing that spindle outward forces are no longer active and cannot maintain the force equilibrium against kinesin– 14-dependent inward forces (Bannigan et al., 2008).