Paenibacillus is a genus of facultative anaerobic, endospore-forming bacteria, originally included within the genus Bacillus and then reclassified as a separate genus in 1993.[4] Bacteria belonging to this genus have been detected in a variety of environments such as: soil, water, rhizosphere, vegetable matter, forage and insect larvae, as well as clinical samples.[5][6][7][8] The name reflects this fact: Latin paene means almost, and so the Paenibacilli are literally almost Bacilli. The genus includes P. larvae, which is known to cause American foulbrood in honeybees, the P. polymyxa, which is capable of fixing nitrogen and therefore is used in agriculture and horticulture, the Paenibacillus sp. JDR-2 which is known to be a rich source of chemical agents for biotechnology applications and pattern forming strains such as P. vortex and P. dendritiformis discovered in the early 90s,[9][10][11][12][13] which are known to develop complex colonies with intricate architectures[14][15][16][17][18] as is illustrated in the pictures.