Membrane permeability was determined according to the method of Liu et al. (2007). Pericarp discs were removed with a cork borer (5 mm in diameter) from the equatorial region of 35 fruit. Fifty discs (about 2 g) were rinsed twice and then soaked in 25 m Lof distilled water at 25◦C for 30 min. The electrolyte leakage (C1)was determined with a conductivity meter (Model 3173, Shanghai Electronics Co., Ltd., China). Another batch of 50 discs was boiled for 15 min in 25 mL of distilled water and then cooled rapidly to25◦C to assess total electrolytes (C2). The relative leakage (%) was expressed as (C1/C2) × 100%.