The question remains whether the new position of Euschemon makes sense in a morphological or other way. Apart from the wingcoupling mechanism, which should be considered a reversal in Euschemon, its new position agrees well with the progressive reduction of the second medial vein (M2) in the hindwing, which is well developed in most Coeliadinae and Euschemon (as in all out-groups), but becomes weaker in the remaining Hesperioidea,particularly in the crown group Hesperiinae, where it may be virtually absent. Therefore, apart from the wing coupling system,the position appears similarly supported by both morphological and molecular evidence. The new position of Euschemon also supports the hypothesis that the early diversification of the Hes-perioidea took place in the eastern part of the Old World tropics in the Cretaceous (de Jong, 2007).