of the distance from the caudal-fin origin beyond the last scale on the caudal base to the margin of the caudal fork (Fig. 2A); an anal-fin origin opposing the 2nd–4th branched dorsal-fin rays; eight scale rows above the lateral line; dark pigment on the dorsal fin concentrated mainly along the margins of the rays (Fig. 3A); 36 or more precaudal lateral-line scales; small dentary tubercles; and, if present, a short, small pair of rostral barbels.