The pH range for growth was investigated
at pH 4.0–11.0 (at intervals of 1.0 pH unit) using the buffer
system described by Xu et al., (2005) and TSB as the basal
medium. The ability to grow under anaerobic conditions was
determined on TSA in an anaerobic jar containing the
anaerobic catalyst Anaerocult A (Merck). Results are given in
the species description