Neighbor-joining (distance-based), maximum parsimony, and maximum-likelihood trees had similar topologies (Fig. 2); haplotypes found in clade 2-1 (identified from nested clade analysis of a 95% parsimony network, see Fig. 3 and below) were the most basal or ancestral, with haplotypes found in nested clades 2-2, 2-3, and 2-4 being more derived (Figs. 2, 3).