Phosphorus and sodium concentrations were the highest (1357–1468 mg/kg dry mass and 1319–1474 mg/kg dry mass, respectively) and modulate pH, followed in descending order by potassium (103–521 mg/kg dry mass), copper (96.1–117 mg/kg dry mass), magnesium (11.4–26.8 mg/kg dry mass), calcium (8.17–8.78 mg/kg dry mass), zinc (1.78–2.08 mg/kg dry mass), iron (0.81–1.20 mg/kg dry mass) and manganese (0.11–0.23 mg/kg dry mass).