(1) recovering stage with small previtellongenic oocytes or primary spermatocytes,
(2) growingิ stage with many large nutritive phagocytes,
(3) mature stage with no or few nutritive phagocytes around oocytes or spermatozoa,
(4) partly spawned stage containing few nutritive phagocytes and loosely packed
gametes,
(5) spent stage with gonads without gametes.
Testes and ovaries were not distinguished for purposes of this study.