Leakage of electrolytes (an indicator of membrane damage) on individual seed basis in 1 mL of
deionized water was determined after 3, 6, 12, 18 and 24 h by measuring the conductivity (μS cm-1) of seed soak water, using ‘Nikon’ digital EC meter. Conductivity was measured at 25oC+1 using 20 seeds each single seed replicate.