Moreover, L. cupreus sp. nov. is distinguished from: (1) the all other species in the complex by an outer metacarpal tubercle divided and smaller than the inner metacarpal tubercle (outer tubercle entire, rounded or somewhat triangular, larger than the inner tubercle in the other species); (2) from L. didymus, L. elenae, L. mystaceus, and L. notoaktites by a smooth dorsal surface of the tibiae (with many small horny spines in these species); and (3) from L. didymus,
L. mystaceus, and L. notoaktites by a tarsus and foot with few tubercles (sole of foot and tarsus with many tubercles in these species).