This may not provide full information, because some of flower-related MADS-box genes are later recruited for fruit development. Thus, duplicate genes with possible overlapping expression patterns in flowers may still show differences in expression during fruit development. The above hypothesis turned out to be true in Magnolia, in which the two AGAMOUS genes have expression patterns which differ qualitatively as the fleshy fruit-like sarcotesta develops. In particular, MgAG2 is expressed at increasing levels and MgAG at decreasing ones.