However, seven species were represented by a single individual (e.g., Canthidium sp. 1,
Canthidium sp. 2, Canthidium sp. 4, Canthonella aff. lenkoi, Coprophanaeus bellicosus, Deltochilum pseudoicarus and Uroxys aff. batesi) (Table S2). Only 16% of fragments contained more than
80% of the estimated true richness (Table 1). Seventy-seven percent of species were recorded in the Coimbra Forest (control area) and 22% of all species were unique for this fragment (Table 1