The leafy liverwort genus Frullania possesses red
coloration (Figure 12), grading into nearly black. Deeply
pigmented species can actually require high light, and
account for the presence of this species at high elevations
above timberline or high in the canopy of the tropics. On
Barro Colorado Island, Panama, this genus grows more
quickly in high light intensities than in the shade, attesting
to its adaptations to high light intensity