Materials and methods
This study was performed on the in-shore reef, Matane Walindi (5° 25′S, 150° 05′E), in Kimbe Bay, Papua New Guinea. Sampling was conducted within 4 m2 quadrats arranged along the reef crest in areas containing at least 6 colonies of each of A. hyacinthus and Pocillopora damicornis. By positioning these quadrats along the reef crest we ensured they were within the home ranges of L. unilineatus and C. baronessa. Along the reef crest these species form a mosaic of overlapping home ranges. The 15 quadrats were divided into 5 groups of 3 adjacent quadrats, and each of these 3 quadrats was then randomly assigned to one of three treatments: (1) bleaching treatment, (2) disturbance control, and (3) unmanipulated control. These quadrats spanned a distance of 110 m along the reef crest, with the minimum distance between quadrats being 5 m.