The plant frequencies of R. acris and L. pratensis were more strongly affected by thefertilizer treatment than by the herbicide treatment, although both treatmentsreduced the plant frequencies significantly. In June 2012, the average decreasein the frequency of R. acris was 22% in the herbicide treated plots (H: mean plantfrequency 45% ± 8 standard error [SE]) and 47% in the fertilizer treated plots (F:mean plant frequency 31% ± 7 SE) compared with the control plots (C: meanplant frequency = 58% ± 4 SE) (Fig. 3C). The effects for L. pratensis were similar,but with a much stronger reduction (nearly twice as large) of the plant frequencyin all treatments. In June 2012, the fertilizer caused an average decrease of 70%(F: mean plant frequency 16% ± 5 SE) in the frequency of L. pratensis, and theherbicide caused an average decrease in the frequency of almost 50% (H: meanplant frequency 28% ± 6 SE) compared with the control plots (C: mean plantfrequency = 53% ± 8 SE (June 2012) (Fig. 3F).