Two genes with a potential role in nutrient mobilization are COPPER CHAPERONE (CCH) and RESPONSIVE-TO-ANTAGONIST1 (RAN1) (Himelblau and Amasino 2000). CCH encodes an intracellular copper chaper- one (Himelblau et al. 1998) while RAN1 encodes a membrane- associated copper-transporting ATPase (Hirayama et al.
1999).