Broad transparent membrane on first gill arch; first ceratobranchial broadly connected to opercular wall; gill
98 HO ET AL. · Zootaxa 3620 (1) © 2013 Magnolia Press
filaments on second to fourth gill arches, two rows of gill filaments on second and third gill arches, single row of
gill filaments on fourth gill arch; those on inner row of third arch and fourth gill arch about one-third length of
those of other arches; inner surface of fourth gill arch completely connected to body. Single row of 17–18 rakers on
1st gill arch, 3–4 on upper limb and 14 on lower limb, 14–15 paired rakers on 2nd arch, 14–15 paired rakers on 3rd
arch and single row of 12 rakers on 4th arch.