England's terrain mostly comprises low hills and plains, especially in central and southern England. However, there are uplands in the north (for example, the mountainous Lake District, Pennines, and Yorkshire Dales) and in the south west (for example, Dartmoor and the Cotswolds). The capital is London, which is the largest metropolitan area in both the United Kingdom and the European Union.[nb 1] England's population of over 53 million comprises 84% of the population of the United Kingdom, largely concentrated around London, the South East, and conurbations in the Midlands, the North West, the North East, and Yorkshire, which each developed as major industrial regions during the 19th century.[9]