During fermentation, the gas produced by yeast causes expan- sion of the gas cells previously incorporated into the batter/dough during the mixing phase (Delcour and Hoseney, 2010). Since the stability and the growth of gas bubbles generated determines the volume of the bread loaf as well as the product texture, any modification during the fermentation phase can alter the crumb structure of the product (He and Hoseney, 1991).