However, there
is an increasing body of literature that indicates that inmany
smaller protected areas, where lions are exposed to various
intense forms of anthropogenic mortality, they do not occur
at predicted densities (Kiffner et al., 2009; Croes et al., 2011;
Groom et al., 2014) and populations comprise smaller prides
and foraging groups (Bauer, 2003; Bauer et al., 2012).