The frond ismultilayered at the centre composed of loosely arranged parenchy-matous cells interspersed with air spaces towards the dorsal sideand of more tightly packed parenchymatous cells with small airspaces towards the ventral surface (Fig. 5a). It tapers to a one cellthick outer rim (Fig. 5a). The cells of the growing pseudoroot arealso housed with considerable amounts of starch grains in theirplastids (Fig. 5a). It is tempting to speculate that these may functionas statoliths for the positive geotropism in the growing phase of thepseudoroot. There is no hint of any vascular tissue differentiation(Fig. 5a).