A recent genetic review of Philippine species (Oliveros & Moyle 2010) found that at least four additional species-level taxa existed there, and the authors did not have access to DNA of all Philippine species, nor did they look at additional taxa on Borneo or Indonesian islands. The "Philippine Bulbul Ixos philippinus" proved to be at least three species, which are better placed in genus Hypsipetes: Philippine Bulbul H. philippinus, with the nominate race on Luzon and race saturatior on Mindanao and nearby islands (i.e., on Bohol, right); Mindoro Bulbul H. mindorensis (Mindoro and Semirara), and Visayan Bulbul H. guimarasensis (Panay, Negros, etc.). Birders had long suspected there were two or more species among these bulbuls, based on plumage and vocal differences. For example, the now-restricted Philippine Bulbul (shown at right) has white shaft streaks to crown and throat, and the throat is rusty, but the newly-split Mindoro Bulbul is dingy without either a rusty throat or shaft streaks.