Boppré, 2005; Figs. 27e31). Vane-Wright and Boppré (2005) further described hitherto overlooked serial pairs of lateral
abdominal androconial organs in Bia that resemble those found in other Brassolini. Vane-Wright and Boppré (2005) further discussed a number of similarities in juvenile stages and lifestyle (unknown to Miller, 1968 and earlier authors) that also support relationship between Bia and other Brassolini. As noted by Vane-Wright and Boppré (2005, p. 237): “The peculiarities of the androconial systems reported here demonstrates that, even without the evidence available from knowledge of DNA sequences and early stage morphology, the clear relationship of Bia totheowl butterflies (Caligo Hübner,1819) and other Brassolini has literally been ‘staring us in the face’ for over 200 years.” The position of Bia as one of the basal lineages in Brassolini has since been corroborated by detailed molecular studies (Peña et al., 2006) and by the combined study of Wahlberg et al. (2009) e even if the circumscription of Satyrinae to include, amongst others, Morphinae presented therein means that Bia strictly speaking is back in Satyrinae.