Inhibition of EPSPS by glyphosate results in the accumulation of shikimic acid and/or certain hydroxybenzoic acids such as protocate- chuic and/or gallic acid in sensitive plant species (Becerril et al., 1989; Lydon and Duke, 1988), and in B. japonicum (Moorman et al., 1992; Hernandez et al., 1999)