Mean (Standard Error; SE) abundance of two species of amphipods (a) and (b) between
sites (S1, S2; tens of metres apart) in each of two locations (L1, L2; 100 m apart) in each of three
mangrove forests (F1, F2, F3; kilometres apart). Note that each species shows significant variation at
each spatial scale, but these differ between the two species. Patterns of variation at the scale of sites
and locations also vary from one mangrove forest to another.