Daniel Hofstadler discussed a model for the development of hyphal networks where the key focus was on tip sensing of
the environment. The model adopted the same core approach of that discussed by Graeme Boswell, but the main emphasis was on modelling tip movement in more detailed, with network formation based closely on the rule-based techniques of, for example, Meškauskas et al. (2004). A key novel feature of the model was tip-to-tip anastomosis that could be achieved by setting an appropriate tropism mediated by the internal supply of nutrients and spatio-temporal gradients of external autoregulators. This has particular relevance to early network development (Glass et al. 2004). Other more complex tip
movements could be obtained by altering rules for tropism.