It is speculated that during ICS formation inflorescence the meristem changes to abnormal inflorescence through genetic
mechanism and growth regulators, ultimately leading to abnormal growth of sepals. Overexpression of AGL24, SOC1, SVP, STMADS16,MPF2 and MPF2-like-A, and silencing of AP1 have been shown to give rise to secondary sepal growth in Arabidopsis