Construction of data matrices. Sequencer chromatograms were edited and individual-gene, individual-taxon sequences were assembled using the PREGAP4 and GAP4 programs in the Staden package33 (version staden_ solaris-1-5-3) or SEQUENCHER (version 4.5; GeneCodes). Multi-sequence alignments were performed manually using the sequence editor Genetic Data Environment34 (version 2.2) and with MAFFT35 (version 6.716b).