This indicates that, in M. grandiflora, the Angiosperm model of fleshy fruit ripening involving the expression of SEPALLATA genes has supplanted the Gymnosperm model involving the expression of AGL6 genes, although a minute trace of the latter model may still occur in the form of low expression of MgAGL6.2 in the very young sarcotesta and even lower in ripening ones. B-sister genes were first found in Gymnosperms (Becker et al. 2002) and their presence subsequently demonstrated also in Angiosperms.